On Planet Maijii, the Tri Squad decide to recruit their New Generation senpai for the Ultra League, and so split up. She blames Tartarus taking Juda and Mold for her loss to the Andro Defence Force, he enslaves her, saying if she does his work she'll see them again, and sends her to retrieve Devil Splinters. On Planet Gua in the Empire Ruins, Tartarus has Reibatos revive Gina Spectre. Also training are Zearth, Nice and Boy, but Zearth and Hice Start arguing and injure eachother, Grigio flies over, heals them, and introductions are made, as Mebius notes her healing could decide the battle.

At the Ultra Coloseum, Grigio starts training with Mebius, who tells her to find her own style or fighting. On Planet K76, back where his training began, Zero has been brought by Joneus, who explains Zero needs to learn to fight purely in instinct to face Tartarus, for this reason he has brought Leo and Seven, who put him back in the Tactor Gear for training. Melos has candidates, Seven 21 and Xenon among them, and Sora is readying an inter-dimensional tracker. At Galaxy Rescue Force HQ, Queen Izana (who is alive, confusing the hevk out of me on the timeline front) consults with Melos and Sora on the preparations to rescue Yullian. Astra has gone to Ebil to try to find a way to The Kingdom by sensing residual energy, while Seven requests help from Leo.

At Inter-Halactic Defence Force HQ, Zoffy and Taro have returned with Leo, relating what King said. Bat Seijin reports to Tartarus that Deabolick and Ultroid Zero Tartarus acquired have been modified. On The Kingdom's Central Planet, Absolution Palace, Tartarus, Diavolo and Titan discuss the Ultra-Ancient power known as the Eternity Core, which they make plans to claim, even if 3 Dark Giants want it themselves. The Destined Crossroads: After the events of The Absolute Conspiracy and the Prologue, Yullian, seeing her cell mates is suffering uses some energy to help him recover, he recalls his Master Alude and probable lover Spica.

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